Carol W. Greider

Carolyn Widney "Carol" Greider (born April 15, 1961) is a molecular biologist at the Johns Hopkins University. She was a co-discoverer of the enzyme telomerase in 1984 while working under Elizabeth Blackburn at the University of California, Berkeley. Greider pioneered research on the structure of telomeres, the ends of the chromosomes. She was awarded the 2009 Nobel Prize for Physiology or Medicine, along with Elizabeth Blackburn and Jack W. Szostak of Boston, Massachusetts, for their discovery of how telomeres are protected from progressive shortening by the enzyme telomerase.